When Pi, an orphaned fish, escapes the fisherman's net to move in with his aunt on an exotic coral reef, he meets his true love, Cordelia. But before Pi can start his life with Cordelia, he must outsmart the bully tiger shark also determined to win her heart.

The fretful Marlin and his young son Nemo become separated from each other in the Great Barrier Reef. Nemo, a clown fish, is unexpectedly taken from his home and thrust into a fish tank in a dentist's office overlooking Sydney Harbor. Buoyed by the companionship of a friendly fish named Dory, Marlin embarks on a dangerous trek and finds himself the unlikely hero.